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1 Postgraduate Research Institute for Sedimentology, Reading University, Whiteknights, Reading RG6 2AB, United Kingdom
2 Department of Earth and Environmental Sciences, Hutchison Hall, University of Rochester, Rochester, New York 14627
The Middle Cambrian Burgess Shale was deposited under variable levels of oxygenation. During periods of oxygenation, low-diversity shell beds were formed and the muds were colonized by infauna. Under these circumstances pyrite was restricted to anoxic microenvironments and formed small (1 mm) discrete aggregated masses. Soft-bodied organisms were preserved under low-oxygen conditions typified by evenly dispersed pyrite, and an absence of trace fossils and shell beds.
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