- Received 5 December 2016.
- Revision received 24 January 2017.
- Accepted 25 January 2017.
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- 1State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Nanjing 210008, China
- 2College of Earth Sciences, University of Chinese Academy of Sciences, Beijing 100049, China
- 3Department of Biological Evolution, Faculty of Biology, Lomonosov Moscow State University, Leninskie gory 1(12), Moscow 119234, Russia
- 4School of GeoSciences, University of Edinburgh, James Hutton Road, Edinburgh EH9 3FE, UK
- 5Siberian Scientific Research Institute of Geology, Geophysics and Mineral Resources, Krasny prospekt 67, Novosibisrk 630091, Russia