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Cover: Fossil shark teeth (Isurus and Carcharocles megalodon). Thousands of teeth like these have been collected from an expansive bonebed in California, called the Sharktooth Hill bonebed. In addition to sharks, the bonebed also preserves a rich assemblage of fossils from extinct marine mammals, sea turtles, bony fish, and seabirds, which accumulated on the seafloor during a pause in deposition that coincided with global temperature maxima and sea-level changes in the middle Miocene. See "Origin of a widespread marine bonebed deposited during the middle Miocene Climatic Optimum" by Pyenson et al., p. 519-522.

Photo courtesy of Randall B. Irmis and Nicholas D. Pyenson

Cover Design by: Heather L. Sutphin